In arid and semiarid areas of the southwestern United States and northwestern México, water availability is the main control on the interactions between the land surface and the atmosphere. Seasonal and interannual variations in water availability regulate the response of water and carbon dioxide fluxes in natural and urban landscapes. However, despite sharing a similar dependance to water availability, landscape characteristics, such as land cover heterogeneity, landscape position, access to groundwater, microclimatic conditions, and vegetation functional traits, among others, can play a fundamental role in modulating the interactions between landscapes and the atmosphere. In this dissertation, I study how different landscape characteristics influence the response of water and carbon dioxide fluxes in arid and semiarid urban and natural settings. The study uses the eddy covariance technique, which calculates the vertical turbulent fluxes within the boundary layer, to quantify water, energy, and carbon dioxide fluxes within local patches. Specifically, the study focused on three main scopes: (1) how vegetation, anthropogenic activity, and water availability influence carbon fluxes in four urban landscapes in Phoenix, Arizona, (2) how access to groundwater and soil-microclimate conditions modulate the flux response of three natural ecosystems in northwestern México during the North American monsoon, and (3) how the seasonal hydrologic partitioning in a watershed with complex terrain regulates the carbon dioxide fluxes of a Chihuahuan Desert shrubland. Results showed a differential response of landscapes according to their land cover composition, access to groundwater or functional traits. In Chapter 2, in urban landscapes with irrigation, vegetation activity can counteract carbon dioxide emissions during the day, but anthropogenic sources from the built environment dominate the carbon dioxide fluxes overall. In Chapter 3, across an elevation-groundwater access gradient, low elevation ecosystems showed intensive water use strategies linked to a dependance to shallow or intermittent access to soil moisture, while a high elevation ecosystem showed extensive water use strategies which depend on a reliable access to groundwater. Finally, in Chapter 4, the mixed shrubland in complex terrain showed an evenly distributed bimodal vegetation productivity which is supported by an abundant water availability during wet seasons and by carry-over moisture in deeper layers of the soil during the dry season. The results from this dissertation highlight how different forms of water availability are responsible for the activity of vegetation which modulates land surface fluxes in arid and semiarid settings. Furthermore, the outcomes of this dissertation help to understand how landscape properties regulate the flux response to water availability in urban and natural areas.

Ecohydrological responses to rainfall in the North American monsoon (NAM) region lead to complex surface-atmosphere interactions. In early summer, it is expected that soil properties and topography act as primary controls in hydrologic processes. Under the presence of strongly dynamic ecosystems, catchment hydrology is expected to vary substantially in comparison to other semiarid areas, affecting our understanding of ecohydrological processes and the parameterization of predictive models. A large impediment toward making progress in this field is the lack of spatially extensive observational data. As a result, it is critical to integrate numerical models, remote sensing observations and ground data to understand and predict ecohydrological dynamics in space and time, including soil moisture, evapotranspiration and runoff generation dynamics. In this thesis, a set of novel ecohydrological simulations that integrate remote sensing and ground observations were conducted at three spatial scales in a semiarid river basin in northern Sonora, Mexico. First, single site simulations spanning several summers were carried out in two contrasting mountain ecosystems to predict evapotranspiration partitioning. Second, a catchment-scale simulation was conducted to evaluate the effects of spatially-variable soil thickness and textural properties on water fluxes and states during one monsoon season. Finally, a river basin modeling effort spanning seven years was applied to understand interannual variability in ecohydrological dynamics. Results indicated that ecohydrological simulations with a dynamic representation of vegetation greening tracked well the seasonal evolution of observed evapotranspiration and soil moisture at two measurement locations. A switch in the dominant component of evapotranspiration from soil evaporation to plant transpiration was observed for each ecosystem, depending on the timing and magnitude of vegetation greening. Furthermore, spatially variable soil thickness affects subsurface flow while soil texture controls patterns of surface soil moisture and evapotranspiration during the transition from dry to wet conditions. Finally, the ratio of transformation of precipitation into evapotranspiration (ET/P) and run off (Q/P) changed in space and time as summer monsoon progresses. The results of this research improve the understanding of the ecohydrology of NAM region, which can be useful for developing sustainable watershed management plans in the face of anticipated land cover and climate changes.